Remy(Patton Oswalt), a provincial rat with a wonderful sense of smell, hates garbage and risks death to enter a human kitchen where he discovers real food and the cooking of five-star chef, Anton Gusteau(Brad Garrett), author of "Anyone Can Cook." On the day Remy learns his hero has died, he is evicted and ends up alone in Paris. By luck, he discovers Gasteau's restaurant, down to three stars and run by a frozen-food-hawking chef. As Remy enters, so does Linguini(Lou Romano), a clumsy youth hired as a garbage boy. To save the soup that Linguini accidentally fouls, Remy throws in some ingredients; the soup is a success and Linguini's career as a chef is born. Can Remy find a way to maintain the fiction and use his gift?