Ben (Sam Elliott) is a cow who for years has been the leader and sober voice of reason among the animals at a farm where the critters are a bit unusual -- they can walk on two legs, talk, swim, and act like humans, though they have the good sense to avoid doing these things while humans are around. Ben has long dreamed that his son Otis ( Kevin James) would someday take over his duties on the farm, but Otis is a carefree and irresponsible type who would rather party with his friends and hang out with his girlfriend, Daisy ( Courteney Cox). Ben and his friend Miles (Danny Glover), a wise and patient mule, wonder if Otis will ever make anything of himself, while Dasiy's best friend, Bessy ( Wanda Sykes), is convinced she can do better. However, one night Otis decides to do something about an obnoxious kid who enjoys tipping his fellow cows, and for the first time in his life he gets a taste of leadership -- and he likes it.